1 | Struwe L,Albert V A.Gentianaceae:systematics and natural history[M].New York:Cambridge University Press,2002:242. |
2 | Ho T N,Liu S W.A worldwide monograph of Swertia and its allies[M].Beijing:Science Press,2015:1-4. |
3 | Von Hagen K B,Kadereit J W.Phylogeny and flower evolution of the Swertiinae(Gentianaceae-Gentianeae):homoplasy and the principle of variable proportions[J].Systematic Botany,2002,27(3):548-572. |
4 | Ho T N,Liu S W.The infrageneric classification of Gentiana(Gentianaceae)[J].Bulletin of the British Museum(Natural History),Botany,1990,20(2):169-192. |
5 | Chen S L,Ho T N,Liu J Q,et al.Embryology of Tripterospermum cordatum(Gentianaceae)[J].Acta Botanica Yunnanica,2000,22(1):53-58. |
6 | Xue C Y,Ho T N,Liu J Q.Embryology of Swertia tetraptera Maxim.(Gentianaceae)and its systematic implication[J].Acta Phytotaxonomica Sinica,1999,37(3):259-263. |
7 | Xue C Y,Ho T N,Liu J Q.Embryology of a Tibetan medicine Halenia elliptica[J].Acta Botanica Yunnanica,1999,21(2):212-217. |
8 | Xue C Y,He T N,Li D Z.Embryology of Swertia cincta(Gentianaceae)and its systematic value[J].Acta Botanica Yunnanica,2002,24(1):75-81. |
9 | Chassot P,Nemomissa S,Yuan Y M,et al.High paraphyly of Swertia L.(Gentianaceae)in the Gentianella-lineage as revealed by nuclear and chloroplast DNA sequence variation[J].Plant Systematics and Evolution,2001,229(1-2):1-21. |
10 | Favre A,Yuan Y M,Küpfer P,et al.Phylogeny of subtribe Gentianinae(Gentianaceae):Biogeographic inferences despite limitations in temporal calibration points[J].Taxon,2010,59(6):1701-1711. |
11 | 郗厚诚,孙瑶,薛春迎.基于ITS和matK序列的獐牙菜亚族(龙胆科龙胆族)分子系统学[J].植物分类与资源学报,2014,36(2):145-156. |
11 | Xi H C,Sun Y,Xue C Y.Molecular phylogeny of Swertiinae(Gentianaceae-Gentianeae)based on sequence data of ITS and matK[J].Plant Diversity and Resources,2014,36(2):145-156. |
12 | 薛春迎,何廷农,李德铢.獐牙菜属植物花蜜腺形态及解剖学[J].云南植物研究,2002,24(3):359-369. |
12 | Xue C Y,He T N,Li D Z.Floral nectaries in Swertia:anatomy and morphology[J].Acta Botanica Yunnanica,2002,24(3):359-369. |
13 | Nemomissa S.Floral character states of the Northeast and Tropical East African Swertia species(Gentianaceae)[J].Nordic Journal of Botany,1997,17(2):145-156. |
14 | Yuan M Y,Küpfer P,Zeltner L.Chromosomal evolution of Gentiana and Jaeschkea(Gentianaceae),with further documentation of chromosome data for 35 species from western China[J].Plant Systematics and Evolution,1998,210(3-4):231-247. |
15 | Nilsson S.Pollen morphological studies in the Gentianaceae—Gentianinae[J].Grana Palynologica,1967,7(1):46-143. |
16 | Kamel E A,Elwan Z A,Loutfy M H A.On the taxonomy of Gentiana L.and Gentianella Moench.(Gentianaceae):evidence from seed coat morphology and seed protein electrophoresis[J].Assiut Univ.J.Bot.,2006,35(2):1-13. |
17 | Kataeva T N,Prokopyev A S,Akinina A A,et al.Seed morphology of some species in the family Gentianaceae[J].Biosciences Biotechnology Research Asia,2015,12(3):2287-2293. |
18 | Whitlock B A,Silver J,Prince J S.Seed coat morphology in Gentianopsis(Gentianaceae)[J].Rhodora,2010,112(949):58-79. |
19 | 葛学军.新疆龙胆科种皮微形态学研究[J].植物研究,1996,16(4):455-458. |
19 | Ge X J.Seed coat micromorphology of Gentianaceae in Xinjiang[J].Bulletin of Botanical Research,1996,16(4):455-458. |
20 | Doyle J J,Doyle J L.A rapid DNA isolation procedure for small quantities of fresh leaf tissue[J].Phytochemical Bulletin,1987,19:11-15. |
21 | Fay M F,Swensen S M,Chase M W.Taxonomic affinities of Medusagyne oppositifolia(Medusagynaceae)[J].Kew Bulletin,1997,52(1):111-120. |
22 | Cuénoud P,Savolainen V,Chatrou L W,et al.Molecular phylogenetics of Caryophyllales based on nuclear 18S rDNA and plastid rbcL,atpB,and matK DNA sequences[J].American Journal of Botany,2002,89(1):132-144. |
23 | Kumar S,Stecher G,Tamura K.MEGA7:molecular evolutionary genetics analysis version 7.0 for bigger datasets[J].Molecular Biology and Evolution,2016,33(7):1870-1874. |
24 | Edgar R C.MUSCLE:multiple sequence alignment with high accuracy and high throughput[J].Nucleic Acids Research,2004,32(5):1792-1797. |
25 | Graham A.Lisianthius pollen from the Eocene of Panama[J].Annals of the Missouri Botanical Garden,1984,71(4):987-993. |
26 | Drummond A J,Suchard M A,Xie D,et al.Bayesian phylogenetics with BEAUti and the BEAST 1.7[J].Molecular Biology and Evolution,2012,29(8):1969-1973. |
27 | Matuszak S,Muellner‐Riehl A N,Sun H,et al.Dispersal routes between biodiversity hotspots in Asia:the case of the mountain genus Tripterospermum(Gentianinae,Gentianaceae)and its close relatives[J].Journal of Biogeography,2016,43(3):580-590. |
28 | Favre A,Michalak I,Chen C H,et al.Out-of-Tibet:the spatio-temporal evolution of Gentiana(Gentianaceae)[J].Journal of Biogeography,2016,43(10):1967-1978. |
29 | Grisebach A.Genera et species Gentianearum adjectis observationibus quibusdam phytogeographicis[M].Stuttgartiae:Bumtibus J.G.Cottan,1839. |
30 | Marquand C V B.The botanical collection made by captain F.Kingdon Ward in the eastern Himalaya and Tibet in 1924-25[J].Botanical Journal of the Linnean Society,1929,48(321):149-229. |
31 | Xue C Y,Ho T N,Li D Z.Embryology of Swertia(Gentianaceae)relative to taxonomy[J].Botanical Journal of the Linnean Society,2007,155(3):383-400. |
32 | Von Hagen K B,Kadereit J W.The phylogeny of Gentianella(Gentianaceae)and its colonization of the southern hemisphere as revealed by nuclear and chloroplast DNA sequence variation[J].Organisms Diversity & Evolution,2001,1(1):61-79. |
33 | 何廷农,薛春迎,王伟.獐牙菜属植物的起源,散布和分布区形成[J].植物分类学报,1994,32(6):525-537. |
33 | Ho T N,Xue C Y,Wang W.The origin,dispersal and formation of the distribution pattern of Swertia L.(Gentianaceae)[J].Acta Phytotaxonomica Sinica,1994,32(6):525-537. |
34 | Card H H.A revision of the genus Frasera[J].Annals of the Missouri Botanical Garden,1931,18(2):245-282. |
35 | Pringle J S.Taxonomic notes on western American Gentianaceae[J].Sida,1990,14(2):179-187. |
36 | 邓涛,吴飞翔,王世骐,等.古近纪/新近纪之交青藏高原陆地生态系统的重大转折[J].科学通报,2019,64(27):2894-2906. |
36 | Deng T,Wu F X,Wang S Q,et al.Significant shift in the terrestrial ecosystem at the Paleogene/Neogene boundary in the Tibetan Plateau[J].Chinese Science Bulletin,2019,64(27):2894-2906. |
37 | Miao Y F,Herrmann M,Wu F L,et al.What controlled Mid–Late Miocene long-term aridification in Central Asia?—Global cooling or Tibetan Plateau uplift:A review[J].Earth-Science Reviews,2012,112(3-4):155-172. |
38 | Li J J,Fang X M.Uplift of the Tibetan Plateau and environmental changes[J].Chinese Science Bulletin,1999,44(23):2117-2124. |
39 | Mulch A,Chamberlain C P.Earth science:the rise and growth of Tibet[J].Nature,2006,439(7077):670-671. |
40 | 刘建全.“整合物种概念”和“分化路上的物种”[J].生物多样性,2016,24(9):1004-1008. |
40 | Liu J Q.“The integrative species concept” and “species on the speciation way”[J].Biodiversity Science,2016,24(9):1004-1008. |
41 | Meyer A,Kocher T D,Basasibwaki P,et al.Monophyletic origin of Lake Victoria cichlid fishes suggested by mitochondrial DNA sequences[J].Nature,1990,347(6293):550-553. |
42 | Chen S Y,Xia T,Wang Y J,et al.Molecular systematics and biogeography of Crawfurdia,Metagentiana and Tripterospermum(Gentianaceae)based on nuclear ribosomal and plastid DNA sequences[J].Annals of Botany,2005,96(3):413-424. |
43 | Wang Y J,Liu J Q,Miehe J.Phylogenetic origins of the himalayan endemic Dolomiaea,Diplazoptilon and Xanthopappus(Asteraceae:Cardueae) based on three DNA regions[J].Annals of Botany,2007,99(2):311-322. |
44 | Zhang X,Deng T,Moore M J,et al.Plastome phylogenomics of Saussurea(Asteraceae:Cardueae)[J].BMC Plant Biology,2019,19:290. |
45 | Lu L M,Mao L F,Yang T,et al.Evolutionary history of the angiosperm flora of China[J].Nature,2018,554(7691):234-238. |