Abstract: The species ivnestigated are Pyrenaria garrettiana Craib, Tutcheria champion! Nakai, T. hirta (H. -M.) Li and T. pingpienensis H. T. Chang. Apart from the number of ovules in each locule, the mega-microsporogenesis and mega-microgatogenesis in all four species are high simi-lar. Embrylogical features shared by these species include: anther tetrasporangiate; anther wall formation of the Basic type; endothecium fibrous; middle layers ephemeral; tapetum glandular; cytokinesis in the microspore mother cell simultaneous; mature pollen grains two-celled; ovules anatropous, bitegmic and tenuinucellate; archesporium one-celled and functioning directly as the megaspore mother cell; embryo sac formation of the Polygonum type; egg cell differentiating dis-tinctly from synergids; integument tapetum and hypostase formed. Of these features, the embryo sac formation of the Polygnoum type forms a main difference from the related genus Camellia whose embryo sac formation conforms to the Allium type.

Key words: Theaceae

摘要： 观察了短叶核果茶(Pyrenaria garrettiana Craib)，石笔木(Tutcheria championi Nakai)，粗毛石笔木(T.hirta(H．－M．)Li)和屏边石笔木(T.pingpienensis H.T．Chang)的大小孢子和雌雄配子体的发生和发育过程，4个种的胚胎学特征高度相似，均为基本型药壁发育，腺质线毡层，同时型小孢子母细胞胞质分裂，四面体形小孢子四分体，二细胞成熟花粉，倒生胚珠，双珠被，薄珠心，单孢原。蓼型胚囊，卵细胞与助细胞区分明显，均具有珠被绒毡层和承珠盘，以蓼型胚囊区别于邻近的山茶属(Camellia)。

关键词: 山茶科, 核果茶属, 石笔术属, 胚胎学