Abstract: The morphology an ddevelopment of female flowers of four species in Hippophae were studied.The result shows that four species are similar,and the differneces among them are only in some quantitative characters.It indicates that properties of the species do not obviously appear as yet at the tissue level during the development of female flowers morphology and embryo sac.But the development rhythm or process in four species is differnet.A new asynchronous meiosis of dyad was discovered in hippopae tibetana Schlecht.The archesporial of H.rhamonides have a long time staying.The development of its stigma and pollen is synchronous but the maturity of the embryo sac which requires warmer condition or certain simulants.It is not reasonable to judge the maturing embryo sac of allicable in H.rhamonides according to the state of stigma opening and its secretion.

Key words: Hippophae, Female flower, Morphology, Development

摘要： 用常规石蜡切片法研究了4种沙棘雌花的发育过程。结果表明4个种极为相似，大部分差别表现在数量特征上，说明种的特征在这一阶段的组织和细胞水平尚未明显反映出来。但是4种沙棘在发育节律上存在差异。西藏沙棘在减数分裂I阶段出现了一种新的二分体的不同步现象；中国沙棘孢原细胞持续时间长，柱头与花粉粒同步发育，而胚囊的形成总是滞后于花粉粒。推断这种现象可能是胚囊的发育需要更温暖的环境条件，或者是花粉管的某种刺激作用；从大孢子母细胞到胚囊成熟进程很快。因此，以柱头张开产生分泌物来判断胚囊成熟对中国沙棘是不适宜的。

关键词: 沙棘, 雌花, 形态, 发育